Professor Harnik is a paleontologist interested in the origin and maintenance of biodiversity in the world's oceans. Harnik received his Ph.D. from the Committee on Evolutionary Biology at the University of Chicago, after which he conducted postdoctoral research at Stanford University and the National Evolutionary Synthesis Center at Duke University. His scholarship has concentrated on the biotic and abiotic drivers of extinction and speciation over geologic time, in particular the evolutionary consequences of rarity. His current research focuses on using paleontological models to predict the response of modern marine ecosystems to current and future environmental change and integrating fossil and molecular data in macroevolutionary analyses.
His full CV can be viewed here.
B.A., Geology, Oberlin College
Ph.D., Evolutionary Biology, University of Chicago
Postdoctoral research, Geological & Environmental Sciences, Stanford University
Postdoctoral reserach, The National Evolutionary Synthesis Center
† Authors contributed equally to this work
* denotes undergraduate student author; ‡ denotes graduate student author
Liow, L.H., T. Reitan, and P.G. Harnik. Submitted to Ecology Letters. Ecological interactions on macroevolutionary time scales: clams and brachiopods are more than ships that pass in the night.
Hoehn, K.B.*, P.G. Harnik, and V.L. Roth. In revision at Methods in Ecology and Evolution. A framework for detecting natural selection on geographic range size at the genus level.
Orzechowski, E.A.*, R. Lockwood, J. Byrnes, S.C. Anderson‡, S. Finnegan, Z.V. Finkel, P.G. Harnik, D.R. Lindberg, L.H. Liow, H.K. Lotze, C.R. McClain, J.L. McGuire, A. O’Dea, J.M. Pandolfi, C. Simpson, and D.P. Tittensor. In revision at Global Change Biology. Determinants of extinction selectivity over the last 500 million years: a meta-analysis of marine bivalves and gastropods.
Finnegan, S.†, S.C. Anderson†‡, P.G. Harnik†, C. Simpson, D.P. Tittensor, J.E. Byrnes, Z.V. Finkel, D.R. Lindberg, L.H. Liow, R. Lockwood, H.K. Lotze, C.M. McClain, J.L. McGuire, A. O’Dea, and J.M. Pandolfi. In press. Paleontological baselines for evaluating extinction risk in the modern oceans. Science.
Harnik, P.G., P.C. Fitzgerald, J.L. Payne, and S.J. Carlson. 2014. Phylogenetic signal in extinction selectivity in Devonian terebratulide brachiopods. Paleobiology 40:675-692. link
Harnik, P.G., C. Simpson, and J.L. Payne. 2012. Long-term differences in extinction risk among the seven forms of rarity. Proceedings of the Royal Society B - Biological Sciences 279:4969-4976. link
Harnik, P.G., H.K. Lotze, S.C. Anderson‡, Z.V. Finkel, S. Finnegan, D.R. Lindberg, L.H. Liow, R. Lockwood, C.M. McClain, J.L. McGuire, A. O’Dea, J.M. Pandolfi, C. Simpson, and D.P. Tittensor. 2012. Extinctions in ancient and modern seas. Trends in Ecology and Evolution 27:608-617. link
Harnik, P.G. 2011. Direct and indirect effects of biological factors on extinction risk in fossil bivalves. Proceedings of the National Academy of Sciences USA 108:13594-13599. link
Harnik, P.G., and R. Lockwood. 2011. Part N, Revised, Volume 1, Chapter 24: Extinction in the marine Bivalvia. Treatise Online 29:1-24 (an online journal presenting chapters of upcoming volumes of the Treatise on Invertebrate Paleontology). PDF
Harnik, P.G., D. Jablonski, A.Z. Krug, and J.W. Valentine. 2010. Genus age, provincial area and the taxonomic structure of marine faunas. Proceedings of the Royal Society B - Biological Sciences 277:3427-3435. link
Harnik, P.G. 2009. Unveiling rare diversity by integrating museum, literature, and field data. Paleobiology 35(2): 190-208. link
Simpson, C., and P.G. Harnik. 2009. Assessing the role of abundance in marine bivalve extinction over the post-Paleozoic. Paleobiology 35(4):631-647. link
Harnik, P.G., and R.M. Ross. 2004. Models of inquiry-based science outreach to urban schools. Journal of Geoscience Education 52(5): 420-428. link
Harnik, P.G., and R.M. Ross. 2003. Developing effective K-16 geoscience research partnerships. Journal of Geoscience Education 51(1): 5-8. link
Harnik, P.G., and R.M. Ross. 2003. Assessing data accuracy when involving students in authentic paleontological research. Journal of Geoscience Education 51(1): 76-84. link
Ross, R.M., Harnik, P.G., Allmon, W.D., Sherpa, J.M., Goldman, A.M., Nester, P.L., and J.J. Chiment. 2003. The Mastodon Matrix Project as an experiment with large-scale collaboration in paleontological research. Journal of Geoscience Education 51(1): 39-47. link
Simonson, B.M., and P.G. Harnik. 2000. Have distal impact ejecta changed through geologic time? Geology 28(11): 975-978. link
Student Theses / Independent Studies/ Summer Students Advised:
2015. Mario Williams, Environmental Science
2014. Rebekah Riemann, Geosciences
2014. Timothy Smith, Geosciences
ENV/GEO 114 - Earth, Environment and Humanity
GEO 221 - History of the Earth
BIO/GEO 275 - Conservation Paleobiology