• hawley pic 2
Visiting Assistant Professor of Psychology



Office: Office #116


  •  2014-current, Visiting Assistant Professor, Department of Psychology, Franklin & Marshall College, Lancaster, PA.
  • 2013-2014, Postdoctoral Fellow, Behavioral Neuroscience, Syracuse University Syracuse, NY.


  • Ph.D, Psychological Sciences & Behavioral Neuroscience, Tulane University, 2013.
  • M.A., Experimental Psychology, Indiana State University, 2008.
  • B.S., Psychology, Magna Cum Laude, SUNY Brockport, 2004.


Stemming from the work I did as a graduate student and as a postdoctoral fellow, my primary research interests revolve around gaining a greater understanding of the biological underpinnings that mediate the relationships between stress and different types of sexual behaviors.  Using an animal model, current research projects include:

-The role of brain-derived neurotrophic factor in male and female rat sexual behaviors and memory for sexual experience.

-The role of estrogen receptors in male and female rat sexual behaviors and memory for sexual experience.



Hawley WR, Witty CF, Daniel JM, Dohanich GP (2015). Choline acetyltransferase in the hippocampus is associated with learning strategy preference in adult male rats. Behavioural Brain Research, 289: 118-24.

Serefoglu EC, Hawley WR, Lasker GF, Grissom EM, Mandava SH, Sikka SC, Dohanich GP, Hellstrom WJ (2014). Effect of botulinum-A toxin injection into bulbospongiosus muscle on ejaculation latency in male rats. Journal of Sexual Medicine, 11: 1657-63.

Hawley WR, Grissom EM, *Moody NM, Dohanich GP, Vasudevan NV (2014). Activation of the GPR30 receptor is sufficient to enhance spatial recognition memory in female rats. Behavioural Brain Research, 262: 68-73.

Ferland CL, Hawley WR, Puckett RE, *Wineberg K, Dohanich GP, Schrader LA (2013). Sirt1 activity in dentate gyrus contributes to chronic stress-induced behavior and ERK1/2 cascade changes in the hippocampus. Biological Psychiatry, 74: 927-935.

Hawley WR, Grissom EM, *Martin RC, *Halmos MB, *Bart CLS, Dohanich GP (2013). Testosterone modulates spatial recognition memory in male rats. Hormones and Behavior, 63: 559-65.

Grissom EM, Hawley WR, *Hodges, KS, *Fawcett-Patel, JM, Dohanich GP (2013). Biological sex and learning strategy preference in prepubertal rats are associated with muscarinic receptor binding in brain areas that modulate place and stimulus-response learning. Hippocampus, 23: 313-22.

Hawley WR, Grissom EM, *Belkin MN, *James TF, Dohanich GP. (2013). Decreased sexual motivation and heightened anxiety in male Long-Evans rats are correlated with the memory for a traumatic event. Archives of Sexual Behavior, 42: 459-68.

Hawley WR, Grissom EM, *Patel JM, *Hodges KS, Dohanich GP. (2013). Reactivation of an aversive memory modulates learning strategy preference. Stress, 16: 73-86.

Grissom EM, Hawley WR, *Bromley-Dulfano SS, *Marino SE, *Stathopoulos NG, Dohanich GP. (2012). Learning strategy is influenced by trait anxiety and early rearing conditions in prepubertal male, but not prepubertal female rats. Neurobiology of Learning and Memory, 2; 174-81.

Hawley WR, Grissom EM, *Barratt HE, *Conrad TS, Dohanich GP. (2012). The effects of biological sex and gonadal hormones on learning strategy in adult rats. Physiology & Behavior, 105: 1014-20.

Hawley W, Grissom E, *Keskitalo L, *Hastings T, Dohanich G. (2011). Sexual motivation and anxiety-like behaviors of male rats after exposure to a trauma followed by situational reminders. Physiology & Behavior, 102: 181-7.

Hawley WR, Grissom EM, Dohanich GP. (2011). The relationships between trait anxiety, place recognition memory, and learning strategy. Behavioural Brain Research, 216: 525-30.

Grissom EM, Brubaker B, Capdevila OS, Hawley WR, Gozal D. (2009). Eye movement during REM sleep in children with attention deficit hyperactivity disorder. Developmental Neuropsychology, 34: 552-9.